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The Many Archaeopteryx by ~Qilong:iconQilong:


©2005-2009 ~Qilong
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Submitted: October 26, 2005
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Varied in form and function, Archaeopteryx stands out as the epitomy of dinobirds, the missing link that's not missing, and only of few species of animals known where every specimen can be recognized by where it's located. It even has a nickname, "Archie" but alas no companion named Jughead. Thank God.

From left to right, largest to smallest:

The Solnhofen Specimen, which has also been called Wellnhoferia grandis -- whose name means "the Large [bird] of Wellnhofer" after Dr. Peter Wellnhofer, who first described the skeleton -- is the largest Archie specimen, and is odd in having an unusual fourth toe.

The Haarlem, or Teylers, Specimen, which was at one point considered to be a pterosaur until 1970, 110 years after it was first found! It was known as Pterodactylus crassipes -- "rough-footed wing-finger" -- until someone noticed the feet were those of a bird, and there were even feathers. It is also the second largest specimen and the least completely preserved.

The Berlin Specimen, which is the holotype, was involved in the controversy surrounding the authenticity of the species, which some claimed was just a dinosaur without feathers, or that the skeleton was real but the feathers weren't, or again that the entire animal was a forgery, which has been extensively disproven.

The "Maxberg" Specimen, which is currently whereabouts unknown, may have been either sold by the private owner after it was taken out of display, stolen, or some other conspiracy related theory ;). The Maxberg, however, does have the oddest wings and legs of any of the Archies.

The Thermopolis Specimen, the newest specimen and the only one not currently residing somewhere in Europe (that we know of). So far, it is one of the best preserved specimens, and has the most detailed palate known to date. The ischium is unusual for all other Archies.

The London Specimen, which has the best preserved braincase of all Archies, is also one of the most studied, based on it's location and preeminence in the bird-origins debate following the publication of Darwin's On the Origin of the Species in 1859.

The München (that's Munich to all you Americans!) Specimen, which is the second smallest Archaeopteryx known and is so far the only recognized other species of Archaeopteryx.

The Eichstätt Specimen, a British researched named Howgate referred this specimen to a new species, Jurapteryx recurva -- "Jurapteryx," for the Jura Mountains of Germany and the Jurassic Period, plus "wing", and "recurva" which refers to the strongly curved foot claws, with relatively larger feet than other specimens -- but which appears to be the smallest A. lithographica known, and perhaps not fully grown.
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Ah, that's coll, a comparison between all "good ol' Archie" :D
Great! Really useful having a visual guide like this to the specimens. I keep getting muddle headed about the details of each...

Only glanced at it so far, but I assume all the details are right.

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"I didn't think today was going to turn out like this."
Hopefully, because the grey areas are either from strict impressions without bone to compensate, or are hesitant fill in's, this won't limit the utility of the panoply. But I paid a good deal to scale and exact measurments, and will soon have a complete table of measurements for the specimens. It would be even more useful had I studied the original material before hand, so I think that actually limits the real utility of this, but for the meanwhile, it's pretty decent....

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"Human beings, who are almost unique in having the ability to learn from the experience of others, are also remarkable for their apparent disinclination to do so." -- Douglas Adams (Last Chance to See)
Thanks for sharing, Jaime, it's cool to have the comparison between all known Archie specimens.
Great work, it's the first time I've seen any illustrative comparison of the individual specimens.

I have to take issue with the taxonomy. The holotype of Archaeopteryx lithographica is the single feather specimen. As this can't be identified to the same genera or species of "Archie" the London specimen has been designated the neotype. The Berlin Specimen was described in 1870, eight years after the London specimen. Buhler & Bock 2002, Journal Fur Ornithologie. v. 143, 3, p. 269.
With respect to Bühler and Bock, I must disagree. While it is true that the feather was described first, when von Wagner mentioned his name, he did in fact refer to the skeleton as he did so. This allowed further workers to immediately regard the skeleton as A. lithographica, while they did so at the time for the feather as well. At this time, the concept of type fixation and holotype was not codified, and people were much more slapdash about it. When the Berlin specimen was described, then named as A. siemensii, and later as Archaeornis siemensii (but now just the former), the London specimen (not the feather) was considered the type specimen. In the 1950's, to avoid confusion over the fixation of specimens, curators at the British Museum appealed to the ICZN and won a fixation of the London specimen as the type. In 1980, a specimen named Pterodactylus crassipes was found in the Teylers Museum of Haarlem, Netherlands, which had apparently been named by von Wagner before A. lithographica, but applied as a specimen of that taxon; this animal had to have its name suppressed for the sake of preserving A. lithographica's position as type species.

So even had von Wagner intended the feather to be the type, the London specimen is fixed as the type by mandate and has largely been accepted as such since. Thus follows all other taxonomists's treatment of the specimens when they review A. lithographica or whatever.

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"Human beings, who are almost unique in having the ability to learn from the experience of others, are also remarkable for their apparent disinclination to do so." -- Douglas Adams (Last Chance to See)
Holy sh!t, that's awesome. You're really good at reconstructions, but tell me, what do you use as referance? pics of the real fossils? other also other sources?

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Olof Moleman AKA Lord Trilobite
Lord of the Paleozoic sea...
No, photos and usually lots of them, with scale attached, are used to describe the detail of the bones. Each one with painstaking tables of measurements accompanying them.

--
"Human beings, who are almost unique in having the ability to learn from the experience of others, are also remarkable for their apparent disinclination to do so." -- Douglas Adams (Last Chance to See)
You forgot to say that the Haarlem specimen is the oldest find of Archaeopteryx (even though it was only later that they realised it)

Btw, I'm also working on a reconstruction of the Haarlem specimen now.

--
Olof Moleman AKA Lord Trilobite
Lord of the Paleozoic sea...

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